Saki Hyuuga and Mai Mishou met at age nine for the first time after they followed two glowing balls that flew towards the Sky Tree, a big tree situated on top of a mountain in their town. Five years later, they met again at the same place and became the new legendary warriors PreCure (Pretty Cure). Flappy and Choppy, spirits from the Land of Fountains, revealed that they were the glowing balls and the girls were chosen to protect the Fountain of Sun hidden in Saki and Mai's world, which they refer to as the Land of Greenery. Saki and Mai are transformed into Cure Bloom and Cure Egret using the spirits. Later they are upgraded to Cure Bright and Cure Windy with help from two additional spirits Mupu and Fupu. The villains of this series are the Dark Fall, who are searching for the Fountain of Sun, the last of the seven fountains that feed the World Tree - the source of all life forms for all worlds. The Leader of the Dark Fall has set his aim on this tree, and it is Pretty Cure's job to protect it.
In the Splash Star movie, Sirlion, a warrior from Dark Fall, opens up a gateway to the Land of Clocks using directions from Mai. His plan to dominate the world is to halt the Eternal (Infinite) Clock and freeze time, cutting off everyone's future. Saki and Mai were already in disagreement after Saki overslept and Mai wandered off into a nearby clock store, causing them to miss the sign-up for the karaoke singing contest. After being thrown into an endless maze, Saki and Mai have to work in unison if they want to solve the puzzle.
